George Ford returns from injury to the England team to face Wales on Saturday – with captain Owen Farrell switching to centre.
The Leicester fly-half has not started for the Red Rose brigade since helping them beat Wales in the Six Nations Championship back in March.
At stake this time is a place in the Nations Cup final and boss Eddie Jones has decided he saw enough from Ford off the bench against Ireland to give him the start.
The match is being played at Parc y Scarlets in Llanelli which Jones has dubbed the “home of Welsh rugby” as that is where the “heart and soul of their game is”.
Wales are given little chance, having lost six of their last seven games, but Jones is wary.
“Twelve months ago Wales were Grand Slam champions and three points from a World Cup final," he said. "So we know what they are capable of.
"We have just got to tidy up our attitude now for the game to make sure we are 100 per cent ready to go right at the start of the game and play with an intensity that Wales have never faced before.”
Following the defensive tour de force against Ireland, particularly from the forwards, Jones leaves his pack unchanged.
Ollie Lawrence misses out at centre due to a hip injury but fit-again duo Luke Cowan-Dickie and Anthony Watson return on the bench.
Jones added: “We’ve picked the best 23 players for the battle and we’re looking forward to it greatly.”
15. Elliot Daly (Saracens, 45 caps)
14. Jonathan Joseph (Bath Rugby, 53 caps)
13. Henry Slade (Exeter Chiefs, 32 caps)
12. Owen Farrell (Saracens, 86 caps)
11. Jonny May (Gloucester Rugby, 59 caps)
10. George Ford (Leicester Tigers, 70 caps)
9. Ben Youngs (Leicester Tigers, 102 caps)
1. Mako Vunipola (Saracens, 62 caps)
2. Jamie George (Saracens, 52 caps)
3. Kyle Sinckler (Bristol Bears, 38 caps)
4. Maro Itoje (Saracens, 41 caps)
5. Joe Launchbury (Wasps, 67 caps)
6. Tom Curry (Sale Sharks, 26 caps)
7. Sam Underhill (Bath Rugby, 20 caps)
8. Billy Vunipola (Saracens, 54 caps)
16. Luke Cowan-Dickie (Exeter Chiefs, 24 caps)
17. Ellis Genge (Leicester Tigers, 21 caps)
18. Will Stuart (Bath Rugby, 6 caps)
19. Jonny Hill (Exeter Chiefs, 2 caps)
20. Ben Earl (Bristol Bears, 6 caps)
21. Jack Willis (Wasps, 1 cap)
22. Dan Robson (Wasps, 5 caps)
23. Anthony Watson (Bath Rugby, 44 caps)
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