Ole Gunnar Solskjaer entered Pep Guardiola's technical area to confront his opposite number during a heated altercation in the Manchester derby.
Solskjaer's United scored early in both halves on their march to a 2-0 victory at the Etihad, with Bruno Fernandes scoring a second-minute penalty and Luke Shaw doubling the advantage in the 50th minute.
City had no response for their visitors' steadfast defence and incisive counter-attacking, and their lacklustre performance at home brought an end to their remarkable run of 21 consecutive victories.
Guardiola cut a frustrated figure on the touchline throughout the encounter and tensions threatened to boil over in the second half as he clashed with United boss Solskjaer.
The dispute started when City players refused to give the ball back to United after Marcus Rashford's injury prompted Dean Henderson to knock the ball out of play.
Rather than give possession back to United, the hosts took the throw-in quickly, leading to a frank exchange of views between Guardiola and Solskjaer.
The two managers initially appeared to make up with a fist-bump only to continue quarrelling moments later.
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The bickering continued for another minute before a second fist-bump restored peace between the two dugouts.
Meanwhile, Rashford – who had been struggling with an ankle injury – was eventually replaced by Mason Greenwood.
United had little trouble seeing out the remainder of the second half, though Guardiola's introduction of Phil Foden for Gabriel Jesus improved his side's attacking threat in the final 20 minutes.
The Red Devils' victory not only ends City's winning run, but also extends Solskjaer's record to three victories in three league games at the Etihad.
United also reclaim second in the Premier League table after Leicester momentarily leapfrogged them with a 2-1 win over Brighton on Saturday, although they remain 11 points behind league-leaders City.
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